This monster appears in the opening, FREEDOM, behind Lester, signifying it as his card.
This monster also appears in the opening, BELIEVE IN NEXUS where he summons his ace card. It also appears in the 5th intro.
In episode 77, Lester obtains this card and its components from the tablet that comes down to Earth after the Three Pure Nobles made note about their plan and Yusei Fudo being the main concern for their mission. Later Lester uses this card during his Turbo Duel against Leo and Luna. He Special Summons this card, "Skiel Top", "Skiel Attack", "Skiel Guard", and "Skiel Carrier" via the effect of "Sky Core". This card's ATK then becomes equal to the combined ATK of every other Skiel monster Lester controls due to its effect. In the next episode, Lester activates "Trap Stun" to negate all other Trap Cards this turn, which prevents Leo from activating his set "Morphtronics, Scramble!" to avoid this card's attack from inflicting Battle Damage to him. This card then attacks Leo directly. On Lester's next turn, he activates this card's effect to equip itself with "Ancient Fairy Dragon" and increase this card's ATK by the ATK of "Ancient Fairy Dragon". Lester then activates "Speed Spell - Summon Close" to forbid Leo and Luna from Special Summoning monsters this turn, which again prevents Leo from activating his set "Morphtronics, Scramble". This card then attacks Leo directly, but Luna activates "Twinkle Wall" to negate the attack and inflict damage to herself equal to half of this card's ATK. Later Leto activates the Defense Position effect of "Morphtronic Vacuumen" to equip this card onto it, but Lester activates "Convert Ghost" to change the target of the effect of "Morphtronic Vacuumen" from this card to the "Absurd Stealer" in his Graveyard, thereby equipping "Absurd Stealer" to "Morphtronic Vacuumen". On Lester's next turn, Leo activates "Power Break" to return all Equip Cards to their owners' Decks and inflict 500 damage to Lester for each Equip Card returned this way. Since two were returned this way, Lester takes 1000 damage. Since "Ancient Fairy Dragon" is no longer equipped onto this card, this card's ATK goes back to 2200. After Lester activates the effect of "Speed World 2" to draw one card, Lester activates "Sky A3" to send "Skiel Attack" to the Graveyard and Special Summon "Skiel Attack 3" in its place. This makes this card's ATK 2400. Lester then Tributes "Skiel Attack 3" in order to Special Summon "Skiel Attack 5". This made this card's ATK 2600. Due to the effect of "Skiel Attack 5", this card can attack directly. This card then attacks Leo directly, defeating him. Since Leo was defeated, all cards on his side of the field are destroyed. Leo would have been killed by the attack, but the Crimson Dragon saves him. Later Luna activates "Ancient Sunshine" to banish "Ancient Fairy Dragon" from her Extra Deck and inflict 2100 damage to Lester, but Lester activates "Infinity Force" to reduce the damage to 0 and destroy all monsters Luna controls. On the last turn of the Duel, this card then attacks directly, defeating Luna.
In episode 103, this card appears in a flashback Leo has when he explains to his friends about the time Lester transferred to his school and when he later Duel against both him and Luna. Leo then noted the damage he felt from that Duel was real.
In episode 108, this card is shown in a flashback Yusei Fudo has when he recognizes the Skiel parts as the parts to the Meklord Emperor that Leo and Luna battled against.
In episode 131, Lester uses this card during his Turbo Duel against Jack Atlas. He Special Summons this card, "Skiel Top", "Skiel Attack", "Skiel Guard", and "Skiel Carrier" via the effect of "Sky Core". Due to this card's effect, this card's ATK becomes equal to the combined ATK of all other Skiel monsters Lester controls. A replay occurs and Jack uses "Red Nova Dragon" to attack this card, but Lester activates the effect of "Skiel Guard" to negate the attack. Jack then activates "Power Pressure" to inflict 1000 damage to Lester. Lester then activates "Infinite Aura", which gains ten Aura Counters (one for each 100 damage Lester took). On Lester's next turn, he activates this card's effect to equip itself with "Red Nova Dragon" and gain ATK equal to the ATK of "Red Nova Dragon", but Jack activates the third effect of "Red Nova Dragon" to banish it until the End Phase and allow Jack to negate one attack this turn. This negates this card's effect as it has now lost its original target. On Lester's End Phase, "Red Nova Dragon" returns to the field. Later "Red Nova Dragon" (whose ATK was 5500 due to its own effect) attacks this card. Lester then activates "Infinite Prison" to discard "Skiel Carrier 5" and set "Sky Core" in his Spell & Trap Card Zone as a Spell Card. "Red Nova Dragon" then destroys this card and wins Jack the Duel. Since this card was destroyed, the other Skiel parts are destroyed too. "Infinite Aura" then gains thirty more Aura Counters since Lester took 3000 damage. Lester then passes the baton to Primo who proceeds in Dueling Jack.
In episode 132, this card appears in a flashback Yusei has when he wonders why the Three Pure Nobles consider it important to use "Infinite Aura". He eventually figures out that they're planning something with it, so he tells Leo to contact Jack and tell him that he needs to destroy "Infinite Aura".
In episode 147, Aporia uses this card during his Ground Duel against Z-one. He Special Summons this card, "Skiel Top", "Skiel Attack", "Skiel Guard", and "Skiel Carrier" via the effect of "Sky Core". Due to this card's effect, its ATK becomes equal to the combined ATK of all other Skiel monsters on Aporia's side of the field. Aporia then activates "High Rate Draw" to destroy all monsters he controls and draw one card for each two monsters destroyed this way rounded down. Since five monsters were destroyed this way, Aporia draws two cards.
This card also appears in Crow's explanation when he explains to Leo that the Meklord Emperors can only be summoned while each of the Meklord Emperor's parts are in the Deck. Since Aporia was milling his Deck for a strategy he had thought out from the beginning of the Duel, Crow explains that it's a good thing that Aporia is using his monsters to help him reduce the number of cards in his Deck.
In episode 149, this monster appears in Z-one's backstory when he explains how he became Yusei and why he wanted to destroy Neo Domino City to change the future. This card is shown on Z-one's computer when he is shown looking up Yusei's history.